Stomach, llama. The first compartment is the fermentation vat. Note the ventral saccules, which have a glandular epithelium. The second compartment seems to function similar to the reticulum of ruminants. Illustration by Dr. Gheorghe Constantinescu. Adapted, with permission, from the Color Atlas of Large Animal Applied Anatomy by Hilary M. Clayton and Peter F. Flood, Mosby-Wolfe, 1996.